Associate Professor
Department of Zoology, University of Florida,
Gainesville, Florida 32611-8525
Telephone: (352) 392-8738
Fax: (352) 392-3704
Internet: mcedward@zoo.ufl.edu

Academic Employment History
• Associate Professor, Department of Zoology, University of Florida (1994-present).
• Visiting Scholar, Department of Biology, Hong Kong University of Science and Technology (1997, 2000)
• Assistant Professor, Department of Zoology, University of Florida (1989-1994).
• Summer Faculty, Friday Harbor Laboratories, University of Washington (1988, 1989, 1991, 1994, 1996, 1999).
• Visiting Scholar, Department of Zoology, University of Washington (1987-1989).
• Summer Faculty, Bamfield Marine Station, Western Canadian Universities Marine Biological Consortium (1987, 1988).
• Killam Post-Doctoral Scholar, Department of Zoology, University of Alberta (1984-1987).

Research Interests
• Marine ecology: Ecology of marine invertebrate larvae.
• Life history evolution: Evolution of development and life cycle patterns.

Education and Degrees
• Ph.D. December 1984, University of Washington, Department of Zoology. On growth and form of echinoplutei. Dissertation advisor: R.R. Strathmann.
• M.A. June 1979, University of South Florida, Department of Biology. Respiratory electron transport system activity during larval development of Lytechinus variegatus Lamarck (Echinodermata:Echinoidea). Thesis advisor: J.M. Lawrence.
• B.A. May 1977, University of New Hampshire, Department of Zoology.

Research Funding (pending)
• National Geographic Society, “The mating system of the starfish Pteraster teselatus: facultative parthenogenesis?”. $19,640, proposal pending
• Environmental Protection Agency, “Assesiing organism to population effects of environmental stress in a sentinel species”. $524,029, 2001-2004. Co-PI with Louis Guillette.
Research Funding (awarded)
• National Science Foundation, "Facultative feeding by planktotrophic larvae of echinoids." $360,000, 1999-2002.
• University of Florida, College of Liberal Arts & Sciences, Research Initiation Project grant. “Effects of the pesticide Alachlor on the embryos and larvae of the sea urchin, Lytechinus variegatus”. $7000, 1997-1998. Co-PI, Dan Brazeau.
• University of Florida, Division of Sponsored Research, Research Development Award. "Graphical computer modeling of embryonic development in sea urchins." $16,000, 1994-1995.
• University of Florida, Division of Sponsored Research, Graduate Research Assistantship Support Program, "The relationship between size, lipid content, and energy content in the eggs of the marine fish, Fundulus heteroclitus" $8942, 1994.
• National Geographic Society, "Evolution of pelagic larval development in pterasterid starfish." $16,424, 1993-1995.
• National Science Foundation, "Effects of an experimental change in egg size on development of planktotrophic larvae of marine invertebrates." $130,000, 1992-1994.
• National Science Foundation, "Effects of an experimental change in egg size on development of planktotrophic larvae of marine invertebrates." Research Experience for Undergraduates Supplement. $5,000, 1993-1994. (Support for 1 student).
• National Science Foundation, "Effects of an experimental change in egg size on development of planktotrophic larvae of marine invertebrates." Research Experience for Undergraduates Supplement. $10,000, 1992-1993. (Support for 2 students).
• University of Florida, Division of Sponsored Research, Research Development Award. "Development and evolution of a novel type of pelagic starfish larva (Echinodermata: Asteroidea)." $20,000, 1990-1991.
• University of Florida, Division of Sponsored Research, Graduate Research Assistantship Support Program, "Morphology and development of a novel type of pelagic starfish larva (Echinodermata: Asteroidea)." $5,419.26, 1989-1990.
• University of Florida, Division of Sponsored Research, Research Support for New Faculty, "Experimental analysis of life history evolution in marine invertebrates." $11,692, 1989.
• University of Alberta, Izaak W. Killam Memorial Post-Doctoral Scholarship. $63,750, 1984-1987.
• University of Alberta, Central Research Fund Operating Grant, "Intraspecific variation in size and organic content of marine invertebrate eggs." $1500, 1985.

Teaching Funding
• University of Florida, Office of Instructional Resources, Multimedia Mini-sabbatical. "Development of computer-based tools for teaching quantitative ecology and evolution in introductory biology." 1996.
• Florida Institute of Oceanography, shiptime awards for “Marine ecology teaching cruise”, on R/V Bellows, $49,350, 1990-1998

Teaching Experience
University of Florida: Cells, Organisms, & Genetics; Evolution, Marine Ecology; Advanced Evolutionary Biology; Integrative Priniples of Biology; Development and Evolution, Seminar in Evolution, Reproduction and Larval Development of Marine Invertebrates; Reproduction and Evolution of Life Histories.
University of Washington (FHL): Marine Invertebrate Zoology; Comparative Invertebrate Embryology; Evolution of Developmental Patterns.
Bamfield Marine Station: Marine Ecology; Larval Biology.

Awards
• Teaching Award (TIP), University of Florida, December 1994.
• Best Paper Award, Coral Reefs, 1990.
• Izaak W. Killam Memorial Post-Doctoral Scholarship, University of Alberta, 1984-1987.

Professional Activities
• Program officer, Division of Invertebrate Zoology, Society for Integrative and Comparative Biology (2000-2002).
• Guest editor for: Canadian Journal of Zoology, special issue on biology of echinoderms (2000); Seminars in Cell and Developmental Biology, special issue, Larval Evolution (2000).
• Graduate Coordinator, Department of Zoology, University of Florida (1996-2000)
• Computer Coordinator, Department of Zoology, University of Florida (1994-1996)
• National Science Foundation, Ocean Sciences grant review panel (1994).
• Reviewer for: National Science Foundation, Natural Sciences and Engineering Research Council (Canada), Natural Environment Research Council (U.K.), National Geographic Society, National Oceanic and Atmospheric Administration, Caribbean Marine Science Center, Science, Evolution, Ecology, American Naturalist, Biological Journal of the Linnean Society, Quarterly Review of Biology, Canadian Journal of Zoology, Seminars in Cell and Developmental Biology, Marine Biology, Marine Ecology Progress Series, Journal of Experimental Marine Biology and Ecology, Biological Bulletin, Pacific Science, Invertebrate Biology, Oceanologica Acta, Invertebrate Reproduction and Development, McGraw-Hill, Cornell University Press, Worth Publishers, Wadsworth Publishing Co., Oxford University Press, Sinauer & Associates, Prentice Hall, University of Queensland (Australia).

Training Experience
• Undergraduate Students (6)
• Bonnie Hall
• Dana Williams
• Jill Borden
• Thom George
• Barry Walling
• Arika Varipongse
• Masters Degree Students: (8)
• Dan Janies
• Ginny Eckert
• Elizabeth Niciu
• Shannon McWeeney
• Marianne Donhue
• Ben Miner
• Adam Reitzel
• Jonathan Cowart
• Ph.D. Degree Students (7)
• Dan Janies
• Joan Herrera
• Andreas Heyland
• Lena Kupriyanova
• Cecelia Miles
• Ben Miner
• Katie Morgan

Publications
• McEdward, L.R., W.B. Jaeckle, and M. Komatsu. (in press). Phylum Echinodermata: Asteroidea. Invited chapter in An Atlas of Invertebrate Larvae, C.M. Young, M.E. Rice, and M. Sewell (eds.), Academic Press.
• McEdward, L.R. (submitted). Calculation and interpretation of the level of egg provisioning in marine invertebrate life cycles. Evolution.
• McEdward, L.R. (submitted). Simulation models of reproductive strategies in marine benthic invertebrates: Fitness differences under fluctuating environmental conditions. Ecology.
• McEdward, L.R. and P.Y Qian. (accepted). Effects of the duration and timing of starvation during larval life on the metamorphosis and juvenile size of the polychaete Hydroides elegans. Journal of Experimental Marine Biology and Ecology.
• McEdward, L.R. and B.G. Miner. (in press). Echinoid larval ecology. Invited review for The Biology and Ecology of Edible Sea Urchins, J.M. Lawrence (ed.), Elsevier, Amsterdam.
• McEdward, L.R. and B.G. Miner. (in press). Larval and life cycle patterns in echinoderms. Invited review for Canadian Journal of Zoology.
• McEdward, L.R. and K.H. Morgan. (in press). Interspecific relationship between egg size and the level of parental investment per offspring in echinoderms. Biological Bulletin.
• McEdward, L.R. 2001. The long and the short of it. Natural History (July, 2001) p.58.
• McEdward, L.R. 2000. Introduction; larval evolution. Seminars in Cell and Developmental Biology 11:383-384. (Invited editorial).
• McEdward, L.R. 2000. Adaptive evolution of larvae and life cycles. Seminars in Cell and Developmental Biology 11:403-409. (Invited review).
• McEdward, L.R. 2000. The origin and evolution of marine invertebrate larvae. Quarterly Review of Biology 75:191-192. Book Review.
• McEdward, L.R. and J.C. Herrera. 1999. Body form and skeletal morphometrics during larval development of the sea urchin Lytechinus variegatus. Journal of Experimental Marine Biology and Ecology 232:151-176.
• McEdward, L.R.. 1997. Reproductive strategies of marine benthic invertebrates revisited: facultative feeding by planktotrophic larvae. Amer. Nat. 150:48-72.
• McEdward, L.R. and D.A. Janies. 1997 . Relationships among development, ecology, and morphology in the evolution of echinoderm larvae and life cycles. Biological Journal of the Linnean Society 60:381-400.
• McEdward, L.R. 1996 Experimental manipulation of parental investment in echinoid echinoderms. American Zoologist. 36:169-179.
• Herrera, J.C., S. McWeeney, and L.R. McEdward. 1996 Diversity of energetic strategies among echinoid larvae and the transition from feeding to nonfeeding development. Oceanologica Acta 19:313-321.
• McEdward, L.R. & M.G. Hadfield. 1996. Flexibility in life cycles. Oceanologica Acta 19:468. (invited conference proceedings paper).
• Nichols, D., L.R. McEdward, & A.B. Smith. 1996. Evolution of life history traits. Oceanologica Acta 19:469-470. (invited conference proceedings paper).
• McEdward, L.R. (ed.) 1995 Ecology of Marine Invertebrate Larvae. CRC Press, Boca Raton.
• McEdward, L.R. 1995 Evolution of pelagic direct development in the starfish Pteraster tesselatus (Asteroidea: Velatida). Biological Journal of the Linnean Society. 154:299-327.
• Janies, D.A. and L.R. McEdward. 1994. A hypothesis for the evolution of the concentricycloid water-vascular system, pp: 246-257, in Reproduction and Development of Marine Invertebrates, W.H. Wilson jr, S.A. Stricker, and G.L. Shinn eds., Johns Hopkins Press
• Janies, D.A. and L.R. McEdward. 1994. Heterotopy, pelagic direct development, and new body plans in velatid asteroids, pp:319-324, in Echinoderms through Time (proceedings of the 8th International Echinoderms Conference, Dijon, France), B. David et al., eds, Balkema Press, Rotterdam, Netherlands.
• McEdward, L. R. and D. A. Janies. 1993. Life cycle evolution in asteroids: What is a larva? Biological Bulletin 184:255-268.
• Janies, D. A. and L. R. McEdward. 1993. Highly derived coelomic and water-vascular morphogenesis in a starfish with pelagic direct development. Biological Bulletin 185:56-76.
• McEdward, L. R. 1992. Morphology and development of a unique type of pelagic larva in the starfish Pteraster tesselatus. Biological Bulletin 182:177-187.
• McEdward, L. R. & F.S. Chia. 1991 Size and energy content of eggs from echinoderms with pelagic lecithotrophic development. Journal of Experimental Marine Biology and Ecology 147:95-102.
• Paulay, G. & L.R. McEdward. 1990 A simulation model of island reef morphology: The effects of sea level fluctuations, growth, subsidence, and erosion. Coral Reefs 9:51-62. Received best paper award from the journal Coral Reefs for 1990.
• Qian, P.Y., L.R. McEdward, & F.S. Chia. 1990 Effects of delayed settlement on survival, growth, and reproduction in the spionid polychaete, Polydora ligni. International Journal of Invertebrate Reproduction and Development 18:147-152.
• McEdward, L.R., S.F. Carson & F.S. Chia. 1988 Energetic content of eggs, larvae, and juveniles of Florometra serratissima and the implications for the evolution of crinoid life histories. International Journal of Invertebrate Reproduction and Development 13:9-21.
• McEdward, L.R. 1988 Experimental embryology as a tool for studying the evolution of echinoderm life histories. In: Echinoderm Phylogeny and Evolutionary Biology, A. Smith & C.R.C. Paul (eds.). Ch. 15, pp:189-196.
• Sinervo, B. & L.R. McEdward. 1988 Developmental consequences of an evolutionary change in egg size: An experimental test. Evolution 42:885-900.
• McEdward, L.R. & R.R. Strathmann. 1987 The body plan of the cyphonautes larva of bryozoans prevents high clearance rates: Comparison with the pluteus and a growth model. Biological Bulletin 172:30-45.
• Emlet, R.B., L.R. McEdward & R.R. Strathmann. 1987 Echinoderm larval ecology viewed from the egg. in: Echinoderm Studies, Vol. 2, pp:55-136, edited by M. Jangoux & J.M. Lawrence. A.A. Balkema.
• McEdward, L.R., & S.F. Carson. 1987 Variation in organic content and its relationship with egg size in the starfish Solaster stimpsoni. Marine Ecology Progress Series 37:159-169.
• McEdward, L.R. & L.K. Coulter. 1987 Egg volume and energetic content are not correlated among sibling offspring of starfish: Implications for life history theory. Evolution 41:914-917.
• McEdward, L.R. 1986 Comparative morphometrics of echinoderm larvae. I. Some relationships between egg size and initial larval form in echinoids. Journal of Experimental Marine Biology and Ecology 96:251-265.
• McEdward, L.R. 1986 Comparative morphometrics of echinoderm larvae. II. Larval size, shape, growth, and the scaling of feeding and metabolism in echinoplutei. Journal of Experimental Marine Biology and Ecology 96:267-286.
• Strathmann. R.R. & L.R. McEdward. 1986 Cyphonautes' ciliary sieve breaks a rule of biological inference. Biological Bulletin 171:694-700.
• McEdward, L.R. 1985 An apparatus for measuring and recording the depth dimension of microscopic organisms. Transactions of the American Microscopical Society 104:194-200.
• McEdward, L.R. 1985 Effects of temperature on the body form, growth, ETS activity, and development rate of an echinopluteus. Journal of Experimental Marine Biology and Ecology 93:169-181.
• Day, R.L. & L.R. McEdward. 1984 Aspects of the physiology and ecology of pelagic larvae of marine benthic invertebrates. pp: 93-120. in: Marine Plankton Life Cycle Strategies, edited by K.A. Steidinger & L.M. Walker. CRC Press.
• McEdward, L.R. 1984 Morphometric and metabolic analysis of the growth and form of an echinopluteus. Journal of Experimental Marine Biology and Ecology 82:259-287.
• McEdward, L.R. & J.M. Lawrence. 1981 Respiratory electron transport system activity during larval development of Lytechinus variegatus Lamarck (Echinodermata: Echinoidea). Comparative Biochemistry and Physiology 70B:653-655.
• Diehl, W.J., L.R. McEdward, E. Proffitt, V. Rosenberg & J.M. Lawrence. 1979 The response of Luidia clathrata (Echinodermata: Asteroidea) to hypoxia. Comparative Biochemistry and Physiology 62A:669-671.

Seminars, symposia and contributed papers (* = presented by co-author)
• 6/2000 Larval Biology, Santa Cruz, CA
• 1/2000 Society of Integrative and Comparative Biology, Atlanta, GA.
• 2/99 American Society of Limnology and Oceanography, Santa Fe, NM
• 1/99 Society of Integrative and Comparative Biology, Denver, CO
• 1/99 Society of Integrative and Comparative Biology(*), Denver, CO
• 7/98 Friday Harbor Laboratories, University of Washington, Friday Harbor, WA
• 1/98 Society of Integrative and Comparative Biology, Boston, MA
• 11/97 F.S. Chia Symposium, Friday Harbor Laboratories, Friday Harbor, WA
• 9/96 Department of Zoology, University of Florida, Gainesville, FL
• 7/96 Friday Harbor Laboratories, University of Washington, Friday Harbor, WA
• 5/96 Department of Fisheries & Aquatic Sciences, University of Florida, Gainesville, FL
• 2/96 Department of Biological Sciences, Florida State University, Tallahassee, FL
• 8/95 Vance Model Symposium, Larval Ecology Meetings, HBOI, Ft. Pierce, FL
• 6/95 Oregon Institute of Marine Biology, University of Oregon, Charleston, OR
• 1/95 American Society of Zoologists(*), St. Louis, MO
• 9/94 Larval Ecology Workshop , Nice & Villefranche-sur-mer, France
• 8/94 Marine Research Institute, Reykjavik, Iceland
• 12/93 Maternal Effects in Life Histories Symposium, ASZ, Los Angeles, CA
• 9/93 8th International Echinoderm Conference(*), Dijon, France
• 12/92 American Society of Zoologists (+3*), Vancouver, BC
• 7/92 North American Echinoderms Conference (+1*), HBOI, Ft. Pierce, FL
• 6/92 Reproduction and Development of Marine Invertebrates Symposium, Friday Harbor, WA
• 4/92 Department of Biology, University of South Florida, Tampa, FL
• 12/91 American Society of Zoologists (+2*), Atlanta, GA
• 7/91 Friday Harbor Laboratories, University of Washington, Friday Harbor, WA
• 3/91 Department of Zoology, Auburn University, Auburn, AL
• 2/91 Whitney Marine Laboratory, University of Florida, St. Augustine, FL
• 5/90 Reproductive Strategies Symposium, Canadian Society of Zoologists, Burnaby, BC
• 4/90 Department of Ecology and Evolution, State University of New York, Stony Brook, NY
• 4/90 Department of Zoology, University of Florida, Gainesville, FL
• 12/89 American Society of Zoologists, Boston, MA
• 2/89 Department of Biology, Williams College, Williamstown, MA
• 2/89 Department of Zoology, University of Florida, Gainesville, FL
• 1/89 Oregon Institute of Marine Biology, University of Oregon, Charleston, OR
• 12/88 American Society of Zoologists, San Francisco, CA
• 7/88 Friday Harbor Laboratories, University of Washington, Friday Harbor, WA
• 1/88 Department of Biology, California State University at Los Angeles, Los Angeles, CA
• 11/87 Department of Biological Sciences, University of California, Santa Cruz, CA
• 11/87 Department of Marine Sciences, University of California, Santa Cruz, CA
• 8/87 International Echinoderms Conference, Victoria, BC
• 12/86 Echinoderm Evolution Symposium, British Museum of Natural History, London
• 10/86 Department of Zoology, University of Alberta, Edmonton, AB
• 9/86 Department of Zoology, University of Alberta, Edmonton, AB
• 5/86 Department of Biology, Marshall University, Huntington, WV
• 12/85 American Society of Zoologists, Baltimore, MD
• 12/84 American Society of Zoologists, Denver, CO
• 12/83 American Society of Zoologists, Philadelphia, PA

revised February 17, 2001